Dineley, D. & Metcalf, S. GCR Editor: D. Palmer. 1999. Fossil Fishes of Great Britain. Geological Conservation Review Series No. 16. JNCC, Peterborough, ISBN 1 86107 470 0. The original source material for these web pages has been made available by the JNCC under the Open Government Licence 3.0. Full details in the JNCC Open Data Policy
Burnham-on-Crouch
Highlights
An extraordinarily rich assemblage of fish taxa is present in the London Clay here. It is almost exclusively of elasmobranchs, and the potential for collecting remains very high.
Introduction
The tidal river cliffs (known as 'The Cliff';
A rich biotic assemblage can be collected loose from the foreshore, and it is possible to bulk sample the clay matrix for further microfossil remains. The fish fauna consists mainly of sharks, and in particular hexanchids, and Burnham is the type locality for several species.
Description
The 2–3 m thick outcrop of London Clay exposed in The Cliff' at Burnham is calculated to be about 26 m below the base of the Claygate Beds, and the strata falls within the top part of Division D (King, 1981), at a palaeontological level defined as Unit P13 by Lake et al. (1986) for the Southend area.
Fauna
Chondrichthyes: Elasmobranchii: Neoselachii: Squalomorphii
Hexanchus agassizi Cappetta, 1976
H. hookeri Ward, 1979
H. collinsonae Ward, 1979
Isistius trituratus (Winkler, 1874)
Notorhynchus serratissimus (Agassiz, 1844)
Squalus minor (Leriche, 1902)
Weltonia burnhamensis Ward, 1979
Chondrichthyes: Elasmobranchii: Neoselachii: Squatinomorphii
Squatina prima (Winkler, 1874)
Chondrichthyes: Elasmobranchii: Neoselachii: Galeomorphii
Alopias crochardi Ward, 1978
Anomotodon sheppeyensis (Casier, 1966)
Carcharias hopei (Agassiz, 1843)
Galeorhinus lefevrei (Daimeries, 1891)
G. minor (Agassiz, 1843)
G. recticonus (Winkler, 1873)
Heterodontus vincenti (Leriche, 1905)
H. woodwardi Casier, 1946
Isurus praecursor (Leriche, 1904)
Isurolamna affinis (Casier, 1946)
Jaekelotodus trigonalis Uaekel, 1895)
Iamna' lerichei Casier, 1946
Megascyliorhinus cooperi Cappetta and Ward, 1977
Mustelus whitei Cappetta, 1976
Odontaspis winkleri (Leriche, 1905)
Otodus obliquus Agassiz, 1836
Palaeohypotodus rutoti (Winkler, 1874)
Palaeorhincodon wardi (Herman, 1975)
Pararhincodon ypresiensis Cappetta, 1976
Physogaleus secundus (Winkler, 1874)
P. tertius (Winkler, 1874)
Scyliorhinus gilberti Casier, 1946
S. burnhamensis Cappetta, 1976
S. casieri Cappetta, 1976
S. pattersoni Cappetta, 1976
S. woodwardi Cappetta, 1976
'Scyliorhinus' minutissimus (Winkler, 1873)
'S.' biauriculatus (Casier, 1950)
Synodontaspis macrotus (Agassiz, 1843)
S. striatus (Winkler, 1874)
Triakis wardi Cappetta, 1976
Xiphodolamia eocaena (Woodward, 1889)
Chondrichthyes: Elasmobranchii: Neoselachii: Batomorphii
Burnhamia daviesi (Woodward, 1889)
Dasyatis davisi Casier, 1966
Myliobatis raouxi Arambourg, 1952
M. dixoni Agassiz, 1843
M. latidens Woodward, 1888
M. toliapicus Agassiz, 1843
Myliobatus sp.
Raja harrisae Ward, 1984
Raja sp.
Interpretation
Much that has been said for the Sheppey site can be repeated here, although the overall number and variety of elasmobranchs is smaller
Comparison with other localities
Burnham-on-Crouch yields a similar fauna, in terms of composition and age, to that from the London Clay series (Division D) of Sheppey (q.v.). However, at Burnham the fish fossils occur free within the clay matrix and can be extracted by bulk sampling methods. Therefore, although whole fossil fishes are not found in nodules, the scattered bones and teeth are recovered in a much better state than similar remains recovered from Sheppey exposures.
Conclusion
Although the London Clay at this site is no more than 3 m thick, the conservation value of the site is derived from its very large fauna of macrofos-sils, both vertebrate and invertebrate. The vertebrate material corresponds to that of the London Clay, division D, at Sheppey. The fact that many taxa are poorly known, despite being first recorded many years ago, should stimulate collecting from a site which is relatively accessible and easy to work.